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They have no tongue, and a complex set of jaws. Chasmfiends, for example, have some flat platelike teeth for crushing and destroying shells and other, smaller mandibles for ripping off flesh or shoving it deeper into the throat.
 
Greatshells live in symbiotic relationships with spren that allow them to grow as large as they do.{{qawob ref|977|158|What are the smoke-y spren that appear around a dead chasmfiend?|date=2012-091227}} These [[Chasmfiend spren|spren]] only appear around the corpses of recently killed greatshells.{{book ref|sa1|15}}
 
== Known Greatshell Species ==
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